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Alan T Whittemore (PhD)
Floral and Nursery Plants Research
Botanist

Phone: (202) 245-4550
Fax: (202) 245-5973
Room 133

U.S. NATIONAL ARBOETUM
3501 NEW YORK AVENUE, NE.
ADMINISTRATION
WASHINGTON, DC 200021958

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Publications (Clicking on the reprint icon Reprint Icon will take you to the publication reprint.)
A new nothospecies in Lagerstroemia (Lythraceae) Reprint Icon - (Peer Reviewed Journal)
Whittemore, A.T., Schori, M. 2022. A new nothospecies in Lagerstroemia (Lythraceae). Phytotaxa. https://doi.org/10.11646/phytotaxa.539.3.10.
The genetic diversity of triploid Celtis pumila and its diploid relatives C. occidentalis and C. laevigata (Cannabaceae) Reprint Icon - (Peer Reviewed Journal)
Hayes, A., Wang, S., Whittemore, A.T., Smith, T. 2022. The genetic diversity of triploid Celtis pumila and its diploid relatives C. occidentalis and C. laevigata (Cannabaceae). Systematic Botany. 47(2):441-451. https://doi.org/10.1600/036364422X16512564801713.
International registration of cultivar names for unassigned woody genera: December 2016-January 2021 Reprint Icon - (Peer Reviewed Journal)
Lura, S.B., Whittemore, A.T. 2021. International registration of cultivar names for unassigned woody genera December 2016-January 2021. HortScience. https://doi.org/10.21273/HORTSCI15914-21.
Phylogeny, biogeography, and classification of the elms (Ulmus) Reprint Icon - (Peer Reviewed Journal)
Whittemore, A.T., Fuller, R.S., Brown, B.H., Hahn, M., Gog, L., Weber, J.A., Hipp, A.L. 2021. Phylogeny, biogeography, and classification of the elms (Ulmus). Systematic Botany. 46(3):711-727. https://doi.org/10.1600/036364421X16312068417039.
Leaf shape and size variation in bur oaks: an empirical study and simulation of sampling strategies Reprint Icon - (Peer Reviewed Journal)
Desmond, S.C., Garner, M., Flannery, S., Whittemore, A.T., Hipp, A.L. 2021. Leaf shape and size variation in bur oaks: an empirical study and simulation of sampling strategies. American Journal of Botany. 108(8):1540-1554. https://doi.org/10.1002/ajb2.1705.
Convergent molecular evolution among ash species resistant to the Emerald Ash Borer Reprint Icon - (Peer Reviewed Journal)
Kelley, L.J., Plumb, W.J., Carey, D.W., Mason, M.E., Cooper, E.D., Crowther, W., Whittemore, A.T., Rossiter, S.J., Koch, J.L., Buggs, R.J. 2020. Convergent molecular evolution among ash species resistant to the Emerald Ash Borer. Nature Ecology and Evolution. 4:1116-1128. https://doi.org/10.1038/s41559-020-1209-3.
Ploidy of seeds from odd-polyploid American elms (Ulmus americana) Reprint Icon - (Peer Reviewed Journal)
Whittemore, A.T., Xia, Z. 2020. Ploidy of seeds from odd-polyploid American elms (Ulmus americana). Journal of the American Society for Horticultural Science. 145:186-192. https://doi.org/10.21273/JASHS04828-19.
Genomic identity of white oak species in the eastern North American syngameon - (Peer Reviewed Journal)
Hipp, A.L., Whittemore, A.T., Garner, M., Hahn, M., Fitzek, E., Guichoux, E., Cavender-Bares, J., Gugger, P.F., Manos, P.S., Pearse, I.S., Cannon, C.H. 2019. Genomic identity of white oak species in the eastern North American syngameon. Annals of the Missouri Botanical Garden. 104:455-477.
Population genetics of Quercus macrocarpa - (Peer Reviewed Journal)
Garner, M., Pham, K., Whittemore, A.T., Cavender-Bares, J., Gagger, P.F., Manos, P.S., Pearse, I.S., Hipp, A.L. 2019. Population genetics of Quercus macrocarpa. International Oaks. 30:131-138.
Berberidaceae - (Peer Reviewed Journal)
Menispermaceae - (Book / Chapter)
Patterns and predictors of fleshy fruit phenology at five international botanical gardens - (Peer Reviewed Journal)
Gallinat, A.S., Primack, R.B., Willis, C.G., Nordt, B., Stevens, A., Fahay, R., Whittemore, A.T., Du, Y., Panchen, Z.A. 2018. Patterns and predictors of fleshy fruit phenology at five international botanical gardens. American Journal of Botany. 105(11):1824-1834.
International registration of cultivar names for unassigned woody genera March 2013 - November 2016 - (Peer Reviewed Journal)
Lura, S.B., Whittemore, A.T. 2018. International registration of cultivar names for unassigned woody genera March 2013 - November 2016. HortScience. 53(8):1232-1237.
What is Sambucus mexicana (Adoxaceae)? - (Peer Reviewed Journal)
Whittemore, A.T. 2018. What is Sambucus mexicana (Adoxaceae)?. Journal of Botanical Research Institute of Texas. 12:69-73.
Ploidy variation in Fraxinus L. (Oleaceae) of eastern North America - (Peer Reviewed Journal)
Whittemore, A.T., Campbell, J.J., Xia, Z., Carlson, C.H., Atha, D., Olsen, R.T. 2018. Ploidy variation in Fraxinus L. (Oleaceae) of eastern North America. International Journal of Plant Sciences. 179(5):377-389. Available: https://www.journals.uchicago.edu/doi/10.1086/696688.
Ranunculaceae - (Book / Chapter)
Genome size variation in elms (Ulmus spp.) and related genera - (Peer Reviewed Journal)
Whittemore, A.T., Xia, Z. 2017. Genome size variation in elms (Ulmus spp.) and related genera. HortScience. 52(4):547-553.
Typification of infrageneric names in Ulmus L. - (Peer Reviewed Journal)
Whittemore, A.T. 2017. Typification of infrageneric names in Ulmus L.. Phytotaxa. 297:291-294.
Adoxaceae - (Book / Chapter)
Ulmaceae for Flora of Missouri - (Book / Chapter)
Whittemore, A.T. 2014. Ulmaceae for Flora of Missouri. Flora of Missouri. 1178-1192.
Leafing out phenology in woody plants of the Northern Hempisphere show phylogenetic, ecological and anatomical patterns - (Peer Reviewed Journal)
Panchen, Z., Primack, R., Nordt, B., Ellwood, E., Stevens, A., Renner, S., Willis, C., Fahey, R., Whittemore, A.T., Du, Y., Davis, C. 2014. Leafing out phenology in woody plants of the Northern Hempisphere show phylogenetic, ecological and anatomical patterns. New Phytologist. DOI: 10.111/nph.12892.
Proposal to conserve the name Celtis glabrata Steven ex Planch. (Cannabaceae) with a conserved type - (Peer Reviewed Journal)
Whittemore, A.T. 2012. Proposal to conserve the name Celtis glabrata Steven ex Planch. (Cannabaceae) with a conserved type. Taxon. 61:254-255.
Ulmus americana is a polyploid complex - (Peer Reviewed Journal)
Whittemore, A.T., Olsen, R.T. 2011. Ulmus americana is a polyploid complex. Journal of Botanical Research Institute of Texas. 98(4):754-760.
Ranunculus - (Book / Chapter)
Whittemore, A.T. 2012. Ranunculus. Jepson Manual of the Higher Plants of California, Ed. 2. pp. 1144-1150.
Validation of the hybrid flowering cherry Prunus xincam Ingram (Rosaceae) - (Peer Reviewed Journal)
Olsen, R.T. and Whittemore, A.T. 2009. Validation of the hybrid flowering cherry Prunus xincam (Rosaceae). Novon. 19:490-493.
Myosurus - (Book / Chapter)
Whittemore, A.T. 2012. Myosurus. Jepson Manual of the Higher Plants of California, Ed. 2. pp. 1142-1144.
Sapindaceae and Acer - (Book / Chapter)
Whittemore, A.T. 2012. Sapindaceae and Acer. Jepson Manual of the Higher Plants of California, Ed. 2. pp. 1230-1231.
Moraceae - (Book / Chapter)
Whittemore, A.T., Mcclintock, E. 2012. Moraceae. Jepson Manual of the Higher Plants of California, Ed. 2. pp. 910-911.
JUGLANDACEAE - (Book / Chapter)
Whittemore, A.T. 2012. JUGLANDACEAE. Jepson Manual of the Higher Plants of California, Ed. 2. pp.832-833.
Pittosporaceae - (Book / Chapter)
Mcclintock, E., Whittemore, A.T. 2012. Pittosporaceae. Jepson Manual of the Higher Plants of California, Ed. 2. pp. 1000-1001.
Cannabaceae (family description and keys) and Celtis - (Peer Reviewed Journal)
Whittemore, A.T. 2012. Cannabaceae (family description and keys) and Celtis. Jepson Manual of the Higher Plants of California, Ed. 2. pp. 598-600.
Ulmaceae - (Peer Reviewed Journal)
Whittemore, A.T. 2012. Ulmaceae. Jepson Manual of the Higher Plants of California, Ed. 2. pp. 1260-1262.
Shepherdia - (Book / Chapter)
Whittemore, A.T. 2012. Shepherdia. Jepson Manual of the Higher Plants of California, Ed. 2. pp. 682-684.
What is Ranunculus gelidus? - (Peer Reviewed Journal)
Whittemore, A.T. 2009. What is Ranunculus gelidus? Journal of Botanical Research Institute of Texas. 3:245-250.
Exotic Species of Celtis (Cannabaceae) in the Flora of North America - (Peer Reviewed Journal)
Whittemore, A.T. 2008. Exotic species of Celtis (Cannabaceae) in the flora of North America. Journal of the Botanical Research Institute of Texas. 2:627-632.
Proposal to Reject the Name Ulmus Campestris L. (Ullmaceae) - (Peer Reviewed Journal)
Whittemore, A.T. 2008. Proposal to reject the name Ulmus campestris (Ulmaceae). Taxon. 57:656-657.
HYBRIDIZATION AND SELF-COMPATIBILITY IN CELTIS: AFLP ANALYSIS OF CONTROLLED CROSSES - (Peer Reviewed Journal)
Whittemore, A.T. and Townsend, A.H. 2007. Hybridization and self-compatibility in Celtis: AFLP analysis of controlled crosses. Journal of the American Society of Horticultural Science. 132:368-373.
Ranunclulaceae - (Book / Chapter)
Parfitt, B.D., Whittemore, A.T. 2013. Ranunclulaceae. Flora of the San Juan Basin. 877-895.
ZINGIBERACEAE - (Book / Chapter)
Whittemore, A.T. Zingiberaceae. The Jepson Manual: Higher Plants of California ed 2. vol. 13.
Hybridization, Invasiveness, and the Impact of Exotic Plants - (Abstract Only)
Hybridization, invasiveness, and the impact of exotic plants. Abstract for Meeting: Invasive Plant Conference: Research, Remove and Renewal, Philadelphia, PA, August 15, 2007.
Hybridization and Introgression in a Sympatric Pair of Oak Species (Fagaceae) Studied with Morphology and Chloroplast and Nuclear SSR Markers - (Abstract Only)
Whittemore, A.T. Hybridization and introgression in a sympatric pair of oak species (Fagaceae) studied with morphology and chloroplast and nuclear SSR markers. Abstract for Meeting: Joint Annual Meeting, American Society of Plant Biologists, American Society of Plant Taxonomists & Botanical Society of America.
The bryophytes of the Departments of Narino and Putumayo, Columbia: II. Hepatics and hornworts - (Peer Reviewed Journal)
Ramirez, P., Whittemore, A.T., Churchill, S.P. 2007. The bryophytes of the Departments of Norina and Putumayo, Columbia: II. Hepatics and hornworts. The Bryologist. 111:363-549.
ITS EVOLUTION IN CELTIS (CANNABACEAE) - (Abstract Only)
Whittemore, A.T. 2006. ITS Evolution in Celtis (Cannabaceae). [abstract]. American Society of Plant Taxonomists Annual Meeting. p. 56.
PITTOSPORACEAE - (Book / Chapter)
Whittemore, A.T. and McClintock, E. 2006. Pittosporaceae. Flora of North America. 13.
NOTES ON SOUTHWESTERN MORACEAE - (Peer Reviewed Journal)
Whittemore, A.T. 2005. Notes on southwestern moraceae. Sida: Contributions to Botany. 22:769-775.
PROPOSAL TO REJECT THE NAME QUERCUS PRINUS (FAGACEAE) - (Peer Reviewed Journal)
Whittemore, A.T., Nixon, K.C. 2005. Proposal to reject the name Quercus prinus L. (Fagaceae). Taxon. 54(1):213-214.
CAPRIFOLIACEAE - (Book / Chapter)
Whittemore, A.T. 2006. Caprifoliaceae. In: Flora of Missouri. 2:765-794.
INTROGRESSION, GENETIC STRUCTURE, AND TAXONOMIC STATUS IN THE CELTIS LAEVIGATA - C. RETICULATA COMPLEX (CELTIDACEAE) - (Peer Reviewed Journal)
Whittemore, A.T. Introgression, Genetic Structure, and Taxonomic Status in The Celtis Laevigata. Systematic Botany.
VALIDATION OF TWO INTERSPECIFIC HYBRID EPITHETS IN ILEX (AQUIFOLIACEAE) - (Peer Reviewed Journal)
Whittemore, A.T. 2005. Validation of Two Name Ilex xaquipernyi J.B. Gable (Aquifoliacceae. Novon. 15:493-494.
IN SEARCH OF NEW PLANTS - (Abstract Only)
In search of new plants
JUGLANDACEAE - (Book / Chapter)
Whittemore, A.T. 2014. JUGLANDACEAE. Flora of Missouri. 282-294.
ULMACEAE - (Book / Chapter)
Fu, L., Yiqun, X., Whittemore, A.T. 2004. Ulmaceae for The Flora of China. Flora of China. v:5, pp:1-19.
SAWTOOTH OAK (QUERCUS ACUTISSIMA, FAGACEAE) IN NORTH AMERICA - (Peer Reviewed Journal)
Whittemore, A.T. 2004. Sawtooth Oak (Quercus Acutissima, Fagaceae) in North America. Sida, Contributions to Botany. v:21, pp:447-454.
BETULACEAE FOR FLORA OF MISSOURI - (Book / Chapter)
Whittemore, A.T. 2006. Betulaceae. In: Flora of Missouri. 2:607-614.