Skip to main content
ARS Home » Northeast Area » Beltsville, Maryland (BARC) » Beltsville Agricultural Research Center » Animal Biosciences & Biotechnology Laboratory » People & Locations » Timothy Ramsay
Timothy G Ramsay
Research Physiologist (Animals)


Publications (Clicking on the reprint icon Reprint Icon will take you to the publication reprint.)
Enteroendocrine peptides, growth, and the microbiome during the porcine weaning transition. - (Peer Reviewed Journal)
Ramsay, T.G., Arfken, A.M., Summers, K.L. 2022. Enteroendocrine peptides, growth, and the microbiome during the porcine weaning transition.. Animal Microbiome. 4:56. https://doi.org/10.1186/s42523-022-00206-8.
Transcriptional reprogramming in rumen epithelium during the developmental transition of pre-ruminant to the ruminant in cattle - (Peer Reviewed Journal)
Baldwin, R.L., Liu, M., Connor, E.E., Ramsay, T.G., Liu, G., Li, C. 2021. Transcriptional reprogramming in rumen epithelium during the developmental transition of pre-ruminant to the ruminant in cattle. Animals. 11(10):2870. https://doi.org/10.3390/ani11102870.
Thermophile lytic enzyme fusion proteins that target Clostridium perfringens - (Peer Reviewed Journal)
Swift, S.M., Reid, K.P., Donovan, D.M., Ramsay, T.G. 2019. Thermophile lytic enzyme fusion proteins that target Clostridium perfringens. Antibiotics. https://doi.org/10.3390/antibiotics8040214.
Yeasts of burden: exploring the mycobiome-bacteriome of the piglet GI tract - (Peer Reviewed Journal)
Arfkin, A.M., Foster Frey, J.A., Ramsay, T.G., Summers, K.L. 2019. Yeasts of burden: exploring the mycobiome-bacteriome of the piglet GI tract. Frontiers in Microbiology. https://doi.org/10.3389/fmicb.2019.02286.
Peripheral histamine and neonatal growth performance in swine - (Peer Reviewed Journal)
Ramsay, T.G., Kahl, S., Long, J.A., Summers, K.L. 2019. Peripheral histamine and neonatal growth performance in swine. Domestic Animal Endocrinology. https://doi.org/10.1016/j.domaniend.2019.06.002.
The piglet mycobiome during the weaning transition: a pilot study - (Peer Reviewed Journal)
Summers, K.L., Foster Frey, J.A., Ramsay, T.G., Arfken, A.M. 2019. The piglet mycobiome during the weaning transition: a ailot study. Journal of Animal Science. https://doi.org/10.1093/jas/skz182.
Conjugated linoleic acid and betaine affect lipolysis in pig adipose tissue explants - (Peer Reviewed Journal)
Ramsay, T.G., Fernández-Fígares, I., Lachica, M., Martínez-Pérez, M. 2019. Conjugated linoleic acid and betaine affect lipolysis in pig adipose tissue explants. Animal-The International Journal of Animal Biosciences. https://doi.org/10.1017/S1751731119001186.
Characterization of two glycosyl hydrolases, putative prophage endolysins, that target Clostridium perfringens - (Peer Reviewed Journal)
Swift, S.M., Donovan, D., Oakley, B.B., Waters, J.J., Rowley, D.T., Ramsay, T.G. 2018. Characterization of two glycosyl hydrolases, putative prophage endolysins, that target Clostridium perfringens. FEMS Microbiology Letters. https://doi.org/10.1093/femsle/fny179.
Orosomucoid binds insulin and IGF1 and reduces hormone stimulated protein synthesis and glucose metabolism in C2C12 myotubes - (Peer Reviewed Journal)
a-1 acid glycoprotein inhibits insulin responses by glucose oxidation, protein synthesis and protein breakdown in mouse C2C12 myotubes - (Peer Reviewed Journal)
Ramsay, T.G., Elsasser, T.H., Caperna, T., Blomberg, L. 2018. a-1 acid glycoprotein inhibits insulin responses by glucose oxidation, protein synthesis and protein breakdown in mouse C2C12 myotubes. Animal. https://doi.org/10.1017/S1751731118001787.
Metabolomic analysis of longissimus from underperforming piglets relative to piglets with normal preweaning growth - (Peer Reviewed Journal)
Ramsay, T.G., Stoll, M.J., Shannon, A.E., Blomberg, L. 2018. Metabolomic analysis of longissimus from underperforming piglets relative to piglets with normal preweaning growth. Journal of Animal and Biotechnology. https://doi.org/10.1186/s40104-018-0251-3.
Use of cross-fostering to enhance growth of pigs that are predicted to grow poorly based on plasma a-1 acid glycoprotein concentration - (Peer Reviewed Journal)
Ramsay, T.G., Stoll, M.J., Schreier, L.L., Shannon, A.E. 2017. Use of cross-fostering to enhance growth of pigs that are predicted to grow poorly based on plasma a-1 acid glycoprotein concentration. Open Journal of Animal Sciences. https://doi.org/10.4236/ojas.2018.81004.
Metabolomic analysis of the longissimus from underperforming piglets suggests impaired carbohydrate metabolism, increased lipid utilization and higher protein breakdown in comparison to piglets with normal preweaning growth - (Abstract Only)
Regulation of fetuin A gene expression in the neonatal pig liver - (Peer Reviewed Journal)
Ramsay, T.G., Stoll, M.J., Blomberg, L., Caperna, T.J. 2017. Regulation of fetuin A gene expression in the neonatal pig liver. Animal-The International Journal of Animal Biosciences. doi:10.1017/S1751731117001410.
A sandwich ELISA for porcine alpha-1acid glycoprotein (pAGP, ORM-1) and further demonstration of its use to evaluate growth potential in newborn pigs - (Peer Reviewed Journal)
Caperna, T.J., Shannon, A.E., Stoll, M.J., Kahl, S., Blomberg, L., Vallet, J.L., Ramsay, T.G. 2017. A sandwich ELISA for porcine alpha-1acid glycoprotein (pAGP, ORM-1) and further demonstration of its use to evaluate growth potential in newborn pigs. Domestic Animal Endocrinology. 60:75-82. doi: 10.1016/j.domaniend.2017.04.001.
Normal birth weight piglets with impaired preweaning growth utilize alternative metabolic pathways in the liver - (Abstract Only)
Regulation of cytokine gene expression by orosomucoid in neonatal swine adipose tissue - (Peer Reviewed Journal)
Ramsay, T.G., Stoll, M.J., Blomberg, L., Caperna, T.J. 2016. Regulation of cytokine gene expression by orosomucoid in neonatal swine adipose tissue. Journal of Animal Science and Biotechnology. 7:25.
Use of plasma orosomucoid in newborn piglets to predict preweaning growth performance and its mechanism of action - (Abstract Only)
Ramsay, T.G., Blomberg, L., Vallet, J.L., Caperna, T.J. 2015. Use of plasma orosomucoid in newborn piglets to predict preweaning growth performance and its mechanism of action. Journal of Animal Science. 94(SUPPLE2):66.
a1-acid glycoprotein inhibits lipogenesis in neonatal swine adipose tissue - (Peer Reviewed Journal)
Ramsay, T.G., Blomberg, L., Caperna, T.J. 2016. a1-acid glycoprotein inhibits lipogenesis in neonatal swine adipose tissue. Animal. 10(5):812-20.
The immune system modulator a1-acid glycoprotein inhibits insulin and IGF1 induced protein synthesis in C2C12 myotubes - (Abstract Only)
Ramsay, T.G., Stoll, M.J., Caperna, T.J., Blomberg, L. 2015. The immune system modulator a1-acid glycoprotein inhibits insulin and IGF1 induced protein synthesis in C2C12 myotubes. Meeting Abstract. 064:40.
Alpha 1-acid glycoprotein has immunomodulatory effects in neonatal swine adipose tissue - (Abstract Only)
Regulation of alpha-1 acid glycoprotein synthesis by porcine hepatocytes in monolayer culture - (Peer Reviewed Journal)
Caperna, T.J., Shannon, A.E., Stoll, M.J., Blomberg, L., Ramsay, T.G. 2015. Regulation of alpha-1 acid glycoprotein synthesis by porcine hepatocytes in monolayer culture. Domestic Animal Endocrinology. 52(7):51-59.
Slow growing pre weaning piglets have altered adipokine gene expression - (Peer Reviewed Journal)
Ramsay, T.G., Stoll, M.J., Caperna, T.J. 2014. Slow growing pre weaning piglets have altered adipokine gene expression. Open Journal of Animal Sciences. 4(7):187-195.
Identification of alpha-1 acid glycoprotein (AGP) as a potential marker of impaired growth in the newborn piglet - (Peer Reviewed Journal)
Caperna, T.J., Shannon, A.E., Blomberg, L., Stoll, M.J., Ramsay, T.G. 2013. Identification of alpha-1 acid glycoprotein (AGP) as a potential marker of impaired growth in the newborn piglet. Reproduction, Fertility and Development. 25:1126-33.
Identification and characterization of a Nuclear Factor Kappa B p65 proteolytic fragment in nuclei of porcine hepatocytes in monolayer culture - (Peer Reviewed Journal)
Caperna, T.J., Shannon, A.E., Garrett, W.M., Ramsay, T.G., Blomberg, L., Elsasser, T.H. 2013. Identification and characterization of a Nuclear Factor Kappa B p65 proteolytic fragment in nuclei of porcine hepatocytes in monolayer culture. Domestic Animal Endocrinology. 45:154-162.
Methyl-ß-cyclodextrin alters adipokine gene expression and glucose metabolism in swine adipose tissue - (Peer Reviewed Journal)
Ramsay, T.G., Blomberg, L., Caperna, T.J. 2014. Methyl-ß-cyclodextrin alters adipokine gene expression and glucose metabolism in swine adipose tissue. Animal-The International Journal of Animal Biosciences. 7(10):1690-1696.
Tumor necrosis a regulation of adipokine gene expression in neonatal adipose tissue - (Peer Reviewed Journal)
Ramsay, T.G., Stoll, M.J., Caperna, T.J., Conde-Aguilera, J.A. 2013. Tumor necrosis a regulation of adipokine gene expression in neonatal adipose tissue. Veterinary Research Communications. 37(1):1-10.
Body composition of piglets exhibiting different growth rates - (Peer Reviewed Journal)
Mitchell, A.D., Ramsay, T.G., Caperna, T.J., Scholz, A.M. 2012. Body composition of piglets exhibiting different growth rates. Archives of Animal Breeding. 55(4):356-363.
Measurement of changes in body composition of piglets from birth to 4 kg using quantitative magnetic resonance (QMR) - (Peer Reviewed Journal)
Mitchell, A.D., Ramsay, T.G., Scholz, A.M. 2012. Measurement of changes in body composition of piglets from birth to 4 kg using quantitative magnetic resonance (QMR). Archives of Animal Breeding. 55:64-71.
Iron Dextran treatment does not induce serum protein carbonyls in the newborn pig - (Peer Reviewed Journal)
Caperna, T.J., Shannon, A.E., Blomberg, L., Garrett, W.M., Ramsay, T.G. 2011. Iron Dextran treatment does not induce serum protein carbonyls in the newborn pig. Cambridge University Press: Animal. 6(1):79-86.
Quantitative magnetic resonance (QMR) measurement of changes in body composition of neonatal pigs - (Proceedings)
Mitchell, A.D., Ramsay, T.G., Taicher, G., Kovner, I. 2010. Quantitative magnetic resonance (QMR) measurement of changes in body composition of neonatal pigs. Symposium on Energy and Protein Metabolism and Nutrition. (G.M. Crovetto, Ed.) EAAP 127:147-148. Wageningen Academic Purlishers. 3rd International Symposium on Engergy and Protein Metabolism, September 6-10, 2010, Parma, Italy.
Identification of Protein Carbonyls in serum of the fetal and neonatal pig. - (Peer Reviewed Journal)
Caperna, T.J., Shannon, A.E., Blomberg, L., Garrett, W.M., Ramsay, T.G. 2010. Identification of protein carbonyls in serum of the fetal and neonatal pig. Comparative Biochemistry and Physiology. 156(3):189-196.
The Effect of Intrauterine Growth Retardation on the Expression of Developmental Factors in Porcine Placenta Subsequent to Placentation - (Peer Reviewed Journal)
Blomberg, L., Schreier, L.L., Guthrie, H.D., Caperna, T.J., Ramsay, T.G., Sample, G.L., Vallet, J. 2010. The effect of intrauterine growth retardation on the expression of developmental factors in porcine placenta subsequent to placentation. Placenta. 31(6):549-52.
Quantitative magnetic resonance (QMR) measurement of changes in body composition of neonatal pigs - (Abstract Only)
Albumin induced cytokine expression in porcine adipose tissue explants - (Abstract Only)
Ramsay, T.G. 2009. Albumin induced cytokine expression in porcine adipose tissue explants. Journal of Animal Science 87, Suppl 2, p. 239.
Analysis of protein oxidation in serum of fetal and newborn piglets and the influence of iron dextran on induction of protein carbonyls. - (Abstract Only)
Caperna, T.J., Shannon, A.E., Ramsay, T.G., Blomberg, L., Garrett, W.M. 2009. Analysis of protein oxidation in serum of fetal and newborn piglets and the influence of iron dextran on induction of protein carbonyls. American Society of Animal Science, Abstract #33536.
IGF-I mediated inhibition of leptin receptor expression in porcine hepatocytes - (Peer Reviewed Journal)
Ramsay, T.G., Richards, M.P., Li, C.J., Caperna, T.J. 2010. IGF-I mediated inhibition of leptin receptor expression in porcine hepatocytes. Comparative Biochemsitry and Physiology. 155(1):43-48.
Impact of runting on adipokine gene expression in neonatal pig adipose tissue - (Peer Reviewed Journal)
Ramsay, T.G., Stoll, M.J., Caperna, T.J. 2010. Adipokine gene transcription level in adipose tissue of runt piglets. Comparative Biochemistry and Physiology. 155(2):97-105.
Uncoupling protein expression in skeletal muscle and adipose tissue in response to in vivo porcine somatotropin treatment - (Abstract Only)
Ramsay, T.G., Mitchell, A.D., Richards, M.P. 2008. Uncoupling protein expression in skeletal muscle and adipose tissue in response to in vivo porcine somatotropin treatment. International Conference on Farm Animal Endocrinology.
Ontogeny of adipokine expression in neonatal pig adipose tissue - (Peer Reviewed Journal)
Ramsay, T.G., Caperna, T.J. 2009. Ontogeny of adipokine expression in neonatal pig adipose tissue. Comparative Biochemistry and Physiology. 152:72-78.
Comparison of Gene Expression in Lean Contemporary and Crossbred Obese Swine - (Peer Reviewed Journal)
Ramsay, T.G., Mitchell, A.D., Richards, M.P. 2007. Comparison of gene expression in lean contemporary and crossbred obese swine. Adipocytes 2(3/4):133-142.
Impact of dietary protein content on uncoupling protein mRNA abundance in swine - (Peer Reviewed Journal)
Ramsay, T.G., Mitchell, A.D. 2008. Impact of dietary protein content on uncoupling protein mRNA abundance in swine. Comparative Biochemistry and Physiology. 149:562-571.
UNCOUPLING PROTEIN EXPRESSION IN SKELETAL MUSCLE AND ADIPOSE TISSUE IN RESPONSE TO IN VIVO PORCINE SOMATOTROPIN TREATMENT - (Peer Reviewed Journal)
Ramsay, T.G., Mitchell, A.D., Richards, M.P. 2008. Uncoupling protein expression in skeletal muscle and adipose tissue in response to in vivo porcine somatotropin treatment. Domestic Animal Endorcinology. 35(2): 130-141.
Beta-adrenergic regulation of uncoupling protein expression in swine. Comparative Physiology and Biochemistry A. 147(2):395-403. - (Peer Reviewed Journal)
Ramsay, T.G., Richards, M.P. 2007. Beta-adrenergic regulation of uncoupling protein expression in swine. Comparative Physiology and Biochemistry A. 147(2):395-403.
LEPTIN IN FARM ANIMALS - ()
Barb, C.R., Hausman, G.J., Ramsay, T.G. 2006. Leptin in farm animals. In: Castracane, D.V., Henson, M.C., editors. Endocrine Update. New York, NY: Springer. p.263-308.
LEPTIN AND LEPTIN RECEPTOR EXPRESSION IN SKELETAL MUSCLE AND ADIPOSE TISSUE IN RESPONSE TO IN VIVO PORCINE SOMATOTROPIN TREATMENT - (Peer Reviewed Journal)
Ramsay, T.G., Richards, M.P. 2005. Leptin and leptin receptor expression in skeletal muscle and adipose tissue in response to in vivo porcine somatotropin treatment. Journal of Animal Science. 83:2501-2508.
LEPTIN AND LEPTIN RECEPTOR EXPRESSION IN SWINE TISSUES IN RESPONSE TO IN VIVO SOMATOTROPIN TREATMENT - (Abstract Only)
Ramsay, T.G., Richards, M.P. 2005. Leptin and leptin receptor expression in swine tissues in response to in vivo somatotropin treatment [abstract]. Journal of Animal Science. 83(Supplement 1):25.
PORCINE PREADIPOCYTE PROLIFERATION AND DIFFERENTIATION: A ROLE FOR LEPTIN? - (Peer Reviewed Journal)
Ramsay, T.G. 2005. Porcine preadipocyte proliferation and differentiation: A role for leptin? Journal of Animal Science. 83(9): 2066-2074.
REGULATION OF UNCOUPLING PROTEINS 2 AND 3 IN PORCINE ADIPOSE TISSUE - (Peer Reviewed Journal)
Ramsay, T.G., Rosebrough, R.W. 2005. Regulation of uncoupling proteins 2 and 3 in porcine adipose tissue. Domestic Animal Endocrinology. v.28:351-356.
HORMONAL REGULATION OF LEPTIN AND LEPTIN RECEPTOR EXPRESSION IN PORCINE SUBCUTANEOUS ADIPOSE TISSUE - (Peer Reviewed Journal)
Ramsay, T.G., Richards, M.P. 2004. Hormonal regulation of leptin and leptin receptor expression in porcine subcutaneous adipose tissue. Journal of Animal Science. v 82:3486-3492.
INTERACTIONS AMONG ENDOCRINE, NUTRITIONAL AND GENETIC FACTORS CONTROLLING METABOLISM IN THE BROILER - (Peer Reviewed Journal)
Rosebrough, R.W., McMurtry, J.P., Richards, M.P., Mitchell, A.D., Ramsay, T.G., Ashwell, C.M. 2005. Interactions among endocrine, nutritional and genetic factors controlling metabolism in the broiler. Avian and Poultry Biology Reviews. 16(2):95-100.
HORMONAL REGULATION OF LEPTIN AND LEPTIN RECEPTOR EXPRESSION IN PORCINE SUBCUTANEOUS ADIPOSE TISSUE - (Abstract Only)
Ramsay, T.G., Richards, M.P. 2004. Hormonal regulation of leptin and leptin receptor expression in porcine subcutaneous adipose tissue [abstract]. Journal of Animal Science. 82 (Suppl. 1):200.
PERIPHERAL LEPTIN ADMINISTRATION ALTERS HORMONE AND METABOLITE LEVELS IN THE YOUNG PIG - (Peer Reviewed Journal)
Ramsay, T.G., Bush, J.A., McMurtry, J.P., Thivierge, M.C., Davis, T.A. 2004. Peripheral leptin administration alters hormone and metabolite levels in the young pig. Comparative Biochemistry and Physiology, Part A. 138:17-25.
PORCINE LEPTIN ALTERS ISOLATED ADIPOCYTE METABOLISM - (Peer Reviewed Journal)
Ramsay, T.G. 2004. Porcine leptin alters isolated adipocyte glucose and fatty acid metabolism. Domestic Animal Endocrinology. 26:11-21.
PORCINE LEPTIN INHIBITS LIPOGENESIS IN PORCINE ADIPOCYTE - (Peer Reviewed Journal)
Ramsay, T.G. 2003. Porcine leptin inhibits lipogenesis in porcine adipocytes.Journal of Animal Science. 81(12):3008-3017.
PORCINE LEPTIN ALTERS FATTY ACID METABOLISM BY SWINE ADIPOCYTES - (Peer Reviewed Journal)
Journal of Animal Science 81:3008-3017, 2003
PORCINE LEPTIN INHIBITS PROTEIN BREAKDOWN IN C2C12 MYOTUBE - (Peer Reviewed Journal)
Ramsay, T.G. 2003. Porcine leptin inhibits protein breakdown and stimulates fatty acid oxidation in C2C12 myotubes. Journal of Animal Science 81(12):3046-3051.
PERIPHERAL LEPTIN ADMINISTRATION ALTERS HORMONE AND METABOLITE LEVELS IN THE YOUNG PIG - (Abstract Only)
Ramsay, T.G., Bush, J.A., McMurtry, J.P., Thivierge, M.C., Davis, R.A. 2003. Peripheral leptin administration alters hormone and metabolite levels in the young pig [abstract]. Journal of Animal Science. v.81(Suppl. 1):95.
HORMONAL REGULATION OF POSTNATAL CHICKEN PREADIPOCYTE DIFFERENTIATION IN VITRO - (Peer Reviewed Journal)
Ramsay, T.G., Rosebrough, R.W. 2003. Hormonal regulation of postnatal chicken preadipocyte differentiation in vitro. Comparative Biochemistry and Physiology Part B. 136:245-253.
LEPTIN REGULATION OF C2C12 MYOTUBE METABOLISM IN VITRO - (Peer Reviewed Journal)
DIETARY CONJUGATED LINOLEIC ACID ALTERS FATTY ACID COMPOSITION OF PIG SKELETAL MUSCLE AND FAT - (Peer Reviewed Journal)
PORCINE LEPTIN ALTERS INSULIN INHIBITION OF LIPOLYSIS IN PORCINE ADIPOCYTES IN VITRO - (Peer Reviewed Journal)
LEPTIN: A REGULATOR OF FEED INTAKE AND PHYSIOLOGY IN SWINE - (Peer Reviewed Journal)
SOMATOTROPIN AND ADIPOSE TISSUE MATABOLISM: SUBSTRATE AND TEMPORAL EFFECTS - (Peer Reviewed Journal)