Technical Abstract: Honeybee foragers actively harvest pollen from cranberry flowers by drumming or stroking the anthers. We show these foragers to be superior cranberry pollinators. More pollen-foragers (86%) delivered pollen in excess of that needed for fruit set, compared with only 50% of nectar- foragers. Stronger colonies fielded more pollen foragers, but syrup supplementation did not enhance pollen foraging. Colonies headed by queens bearing a heritable pollen-hoarding trait harvested significantly more pollen (and so presumably field more pollen- foragers) than standard colonies. Fielding more pollen-foragers translates into delivering more effective cranberry pollinators for a honey bee colony of a given size. Genetic polymorphisms of flowering plants are known to influence foraging behaviors of pollinators; however, no heritable polymorphism of a pollinator is known to influence its efficacy as a pollinator. Cranberry flowers require bee visitation for pollination. Honey bees visit cranberry flowers for nectar but rarely pollen if alternative floral species exist nearby. Floral stigmas of managed cranberries visited once by pollen-foraging honey bees received an average 4-fold more pollen than flowers visited by mere nectar foragers, even when the latter made stigmatic contact.