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United States Department of Agriculture

Agricultural Research Service

Thomas Tworkoski (Tom)

Plant Physiologist


Publications (Clicking on the reprint icon Reprint Icon will take you to the publication reprint.)
The molecular basis for tree growth habit in Prunus persica (peach) - (Abstract Only)
Dardick, C.D., Hollender, C.A., Scorza, R., Srinivasan, C., Callahan, A.M., Zhebentyayeva, T., Ruiz, K.B., Whitaker, M., Horn, R., Abbott, A., Tworkoski, T. 2014. The molecular basis for tree growth habit in Prunus persica (peach)[abstract]. 7th International Rosaceae Genomics Conference. p. 15.
Genetic interactions underlying tree branch orientation - (Abstract Only)
Hollender, C.A., Scorza, R., Callahan, A.M., Tworkoski, T., Dardick, C.D. 2014. Genetic interactions underlying tree branch orientation [abstract]. American Society for Plant Biology. Paper No. P-12023-B.
Interaction of scions and size-controlling rootstocks of young apple trees - (Abstract Only) - (30-Aug-13)
Effects of size-controlling rootstocks on growth and yield of peach trees with upright and spreading growth habits - (Peer Reviewed Journal)
Tworkoski, T., Miller, S.S., Scorza, R. 2013. Effects of size-controlling rootstocks on growth and yield of peach trees with upright and spreading growth habits. Journal of American Pomological Society. 67(2):89-94.
Rotating cross-arm trellis technology for blackberry production - (Peer Reviewed Journal)
Takeda, F., Glenn, D.M., Tworkoski, T. 2013. Rotating cross-arm trellis technology for blackberry production. Journal of Berry Research. 3(1):25-40.
Weed suppression by grasses for orchard floor management - (Peer Reviewed Journal)
Tworkoski, T., Glenn, D.M. 2012. Weed suppression by grasses for orchard floor management. Weed Technology. 26(3):559-565.
Physiological processes of size-controlling rootstocks in apple - (Proceedings)
Tworkoski, T., Fazio, G. 2012. Physiological processes of size-controlling rootstocks in apple. In: Proceedings of the 87th Annual Cumberland-Shenandoah Fruit Workers Conference, December 1-2,2011, Winchester, VA. p. 11.
Long-term effects of sod competition on peach production for standard and pillar growth types on yield and economic parameters - (Peer Reviewed Journal)
Glenn, D.M., Tworkoski, T., Scorza, R., Miller, S.S. 2011. Long-term effects of sod competition on peach production for standard and pillar growth types on yield and economic parameters. HortTechnology. 21(6):720-725.
Gene expression and hormone levels in different growth habits of peach and apple - (Proceedings)
Tworkoski, T., Webb, K.K., Callahan, A.M., Scorza, R. 2012. Gene expression and hormone levels in different growth habits of peach and apple. In: Proceedings of the Plant Growth Regulator Society of America, July 24-27, 2011, Chicago, Illinois. p. 145.
Global changes in expression of grapefruit peel tissue in response to the yeast biocontrol agent, Metschnikowia fructicola - (Peer Reviewed Journal)
Hershkovitz, V., Droby, S., Wisniewski, M.E., Liu, J., Ben-Dayan, C., Raphael, G., Pasmanik-Chor, M., Aly, R., Tworkoski, T. 2012. Global changes in expression of grapefruit peel tissue in response to the yeast biocontrol agent, Metschnikowia fructicola. Molecular Plant Pathology. 13(4):338-349.
Application of BABA and s-ABA for drought resistance in apple - (Peer Reviewed Journal)
Tworkoski, T., Wisniewski, M.E., Artlip, T.S. 2011. Application of BABA and s-ABA for drought resistance in apple. Journal of Applied Horticulture. 13(2):85-90.
Blackberry propagation by non-leafy floricane cuttings - (Peer Reviewed Journal)
Takeda, F., Tworkoski, T., Finn, C.E. 2011. Blackberry propagation by non-leafy floricane cuttings. HortTechnology. 21(2):236-239.
Effect of heat shock treatment on stress tolerance and biocontrol efficacy of Metschnikowia fructicola - (Peer Reviewed Journal)
Liu, J., Wisniewski, M.E., Droby, S., Tian, S., Tworkoski, T. 2011. Effect of heat shock treatment on stress tolerance and biocontrol efficacy of Metschnikowia fructicola. FEMS Microbiology Ecology. 76:145-155.
Grass competition may benefit high density peach orchards - (Proceedings)
Tworkoski, T., Glenn, D.M. 2010. Grass competition may benefit high density peach orchards. Annual Cumberland Shenandoah Fruit Workers Conference, November 18-19, 2010, Winchester, VA. p. 45-49.
Interaction of BABA and ABA for drought resistance in apple - (Proceedings)
Tworkoski, T., Wisniewski, M.E., Artlip, T.S. 2011. Interaction of BABA and ABA for drought resistance in apple. Plant Growth Regulator Society of America Meeting. p. 142.
Long-term effects of managed grass competition and two pruning methods on growth and yield of peach trees Reprint Icon - (Peer Reviewed Journal)
Tworkoski, T., Glenn, D.M. 2010. Long-term effects of managed grass competition and two pruning methods on growth and yield of peach trees. Scientia Horticulturae 126:130-137.
Blossom thinning in apple and peach with an essential oil - (Peer Reviewed Journal)
Miller, S.S., Tworkoski, T. 2010. Blossom thinning in apple and peach with an essential oil. HortScience. 45(8):1218-1225.
Eugenol bloom thinner on peach - (Proceedings)
Miller, S.S., Tworkoski, T., Hott, C.F. 2010. Eugenol bloom thinner on peach. Annual Cumberland Shenandoah Fruit Workers Conference.
Weed suppression by grasses for orchard floor management - (Proceedings)
Tworkoski, T., Glenn, D.M. 2010. Weed suppression by grasses for orchard floor management. Annual Cumberland Shenandoah Fruit Workers Conference.
Leaf N and P in different growth habits of peach: effects of root system morphology and transpiration - (Peer Reviewed Journal)
Tworkoski, T., Scorza, R., Glenn, D.M. 2009. Leaf N and P in different growth habits of peach: effects of root system morphology and transpiration. Journal of Applied Horticulture. 11(2):95-98.
Hormonal and anatomical effects of apple rootstocks - (Proceedings)
Tworkoski, T., Fazio, G. 2010. Hormonal and anatomical effects of apple rootstocks. Proceedings of the 36th annual meeting of the Plant Growth Regulation Society of America, (PGRSA, LaGrange, GA). August 2-6, 2009, Asheville, North Carolina, p. 161.
Effects of peach tree root system morphology and transpiration on leaf nitrogen and phosphorus - (Proceedings)
Tworkoski, T., Scorza, R., Glenn, D.M. 2009. Effects of peach tree root system morphology and transpiration on leaf nitrogen and phosphorus. Annual Cumberland Shenandoah Fruit Workers Conference. Pg 175.
Physiological and morphological effects of size-controlling rootstocks on ‘Fuji’ scion - (Peer Reviewed Journal)
Tworkoski, T., Fazio, G. 2011. Physiological and morphological effects of size-controlling rootstocks on ‘Fuji’ scion. Acta Horticulturae. 903:865-872.
Response of Young Apple Trees to Grass and Irrigation - (Peer Reviewed Journal)
Tworkoski, T., Glenn, D.M. 2008. Response of Young Apple Trees to Grass and Irrigation. International Journal of Fruit Science. 8:89-108.
Physiological and morphological effects of size-controlling rootstocks on 'Fuji' scion - (Abstract Only)
Tworkoski, T., Fazio, G. 2008. Physiological and morphological effects of size-controlling rootstocks on 'Fuji' scion. International Symposium on Integrating Canopy, Rootstock and Environmental Physiology in Orchard Systems in Geneva, NY, August 4-8, 2008.
Response of Young Apple Trees to Grass and Irrigation - (Proceedings)
Tworkoski, T., Glenn, D.M. 2008. Response of young apple trees to grass and irrigation. Annual Cumberland Shenandoah Fruit Workers Conference. November 15-1, 2007, Winchester, VA (Daniel Ward, ed) p.75.
Rooting Response of Shoot Cuttings from Three Peach Growth Habits - (Peer Reviewed Journal)
Tworkoski, T., Takeda, F. 2007. Rooting response of shoot cuttings from three peach growth habits. Scientia Horticultureae. 115:98-100.
Endogenous hormone concentrations and bud break response to exogenous BA in shoots of apple trees with two growth habits grown on three rootstocks - (Peer Reviewed Journal)
Tworkoski, T., Miller, S.S. 2007. Endogenous hormone concentrations and bud break response to exogenous BA in shoots of apple trees with two growth habits grown on three rootstocks. Journal of Horticultural Science and Biotechnology. 82(6):960-966.
Blossom thinning in apple and peach with an essential oil - (Proceedings)
Miller, S.S., Tworkoski, T. 2007. Blossom thinning in apple and peach with an essential oil. Annual Cumberland Shenandoah Fruit Workers Conference.
Endogenous Hormone Concentrations and Bud Break Response to Exogenous BA in Shoots of Apple Trees with Two Growth Habits Grown on Three Rootstocks - (Proceedings)
Tworkoski, T., Miller, S.S. 2007. Endogenous Hormone Concentrations and Bud Break Response to Exogenous BA in Shoots of Apple Trees with Two Growth Habits Grown on Three Rootstocks. Annual Cumberland Shenandoah Fruit Workers Conference. P. 114.
Endogenous Hormone Concentrations and Bud Break Response to Exogenous Ba in Shoots of Apple Trees with Two Growth Habits Grown on Three Rootstocks - (Abstract Only)
Tworkoski, T., Miller, S.S. 2007. Endogenous hormone concentrations and bud break response to exogenous ba in shoots of apple trees with two growth habits grown on three rootstocks. Proceedings of Plant Growth Regulation Society of America. Pg. 225.
Rootstock Effect on Growth of Apple Scions with Different Growth Habits - (Peer Reviewed Journal)
Tworkoski, T., Miller, S.S. 2007. Rootstock effect on growth of apple scions with different growth habits. Scientia Horticulturae 111 p. 335-343.
Endogenous Hormone Concentrations and Bud Break Response to Exogenous Ba in Shoots of Apple Trees with Two Growth Habits Grown on Three Rootstocks - (Abstract Only)
Tworkoski, T., Miller, S.S. 2006. Endogenous hormone concentrations and bud break response to exogenous ba in shoots of apple trees with two growth habits grown on three rootstocks. Plant Growth Regulation Society of America Quarterly. 34(2):52 (Abstract 68).
Enhancing Biological Control in Orchards by Increasing Food Web Biodiversity - (Peer Reviewed Journal)
Brown, M.W., Tworkoski, T. 2006. Enhancing biological control in orchards by increasing food web biodiversity. J. Fruit and Ornamental Plant Research. Vol. 14 (Suppl. 3) 2006., 19-27.
Relationship of Pruning and Growth Morphology with Hormone Ratios in Shoots of Pillar and Standard Peach Trees - (Peer Reviewed Journal)
Tworkoski, T., S. Miller, and R. Scorza. 2006. Relationship of pruning and growth morphology with hormone ratios in shoots of pillar and standard peach trees. J. Plant Growth Regul. 25:1-11.
Rootstock Effect on Growth of Apple Scion with Different Growth Habits - (Proceedings)
Tworkoski, T., Miller, S.S. 2006. Rootstock effect on growth of apple scion with different growth habits. Meeting Abstract. 2005 Cumberland-Shenandoah Fruit Workers Conference 81st Annual Meeting. Volume 31:193.
Rootstock Effect on Growth of Apple Scions with Excurrent and Decurrent Growth Habits - (Proceedings)
Tworkoski, T., Miller, S.S. 2006. Rootstock effect on growth of apple scions with excurrent and decurrent growth habits. Meeting Abstract. Proceedings of the 32nd Annual Meeting of the Plant Growth Regulation Society of Amerida 32:142. July 24-27, 2005, Newport Beach, California.
Differential Regulation of Two Dehydrin Genes from Peach (Prunus Persica) by Photoperiod, Low Temperature and Water Deficit - (Peer Reviewed Journal)
Wisniewski, M.E., Bassett, C.L., Renaut, J., Farrell, Jr., R., Tworkoski, T., Artlip, T.S. 2006. Differential regulation of two dehydrin genes from peach (prunus persica) by photoperiod, low temperature and water deficit. Tree Physiology. 26:575-584.
Rootstock Effect on Growth of Apple Scion with Different Growth Habits - (Abstract Only)
Tworkoski, T., Miller, S.S. 2005. Rootstock effect on growth of apple scion with different growth habits. Plant Growth Regulator Society of America Meeting. July 24-27, 2005. Newport Beach, CA. ABSTRACT NO. 46.
Developing Peach Cultivars with Novel Tree Growth Habits - (Proceedings)
Scorza, R., Okie, W.R., Miller, S.S., Glenn, D.M., Tworkoski, T. 2006. Developing peach cultivars with novel tree growth habits. Acta Horticulturae. 713:61-64.
Orchard Floor Management Systems - (Book / Chapter)
Tworkoski, T., Glenn, D.M. 2008. Orchard floor management systems. In: Layne, D.R., Bassi, D., editors. In the Peach, Botany, Production, and Uses. Cambridge, MA: CBI. p. 332-350.
Morphological and Hormonal Relationships in Shoots of Pillar and Standard Peach Trees - (Proceedings) - (21-Jan-05)
Developing Peach Cultivars with Novel Tree Growth Habits - (Abstract Only)
Scorza, R., Okie, W.R., Miller, S.S., Tworkoski, T., Glenn, D.M. 2005. Developing peach cultivars with novel tree growth habits. Acta Horticulturae. 16p.
Characteristics and Management of Peach and Apple Tree Crowns - (Proceedings)
Tworkoski, T. 2005. Characteristics and management of peach and apple tree crowns. Proceedings of Plant Growth Regulation Society of America. Vol.32
Effects of Pruning on Auxin and Cytokinin Levels and Subsequent Shoot Regrowth among Different Growth Habits of Peach - (Proceedings)
Tworkoski, T., Miller, S.S., Scorza, R. 2005. Effects of pruning on auxin and cytokinin levels and subsequent shoot regrowth among different growth habits of peach. Proceedings of Plant Growth Regulation Society of America. v. 32. p. 46.
Weed Control with Hydrophobic and Hydrous Kaolin Clay Particle Mulches - (Peer Reviewed Journal)
Takeda, F., Glenn, D.M., Tworkoski, T. 2005. New strategies for weed control in blackberry plantings. Hortscience. Vol 40(3): 714-719.
Characteristics and Management of Peach and Apple Tree Crowns - (Abstract Only)
Tworkoski, T. 2004. Characteristics and management of peach and apple tree crowns. Plant Growth Regulator Society of America Meeting. 32:31
Effects of Pruning on Auxin and Cytokinin Levels and Subsequent Shoot Regrowth of Different Growth Habits of Peach - (Abstract Only)
Tworkoski, T., Miller, S.S., Scorza, R. 2004. Effects of pruning on auxin and cytokinin levels and subsequent shoot regrowth of different growth habits of peach. Meeting Abstract. 32:48
Pest Management Benefits of Compost Mulch in Orchards - (Peer Reviewed Journal)
Brown, M.W., Tworkoski, T. 2004. Pest management benefits of compost mulch in orchards. Agricultural Ecosystems and Environment. (2004) 103:465-472
"effects of a Root Barrier and Localized Fertilizer Application on Root Growth of Young Peach (Prunus PERSICA)TREES." - (Peer Reviewed Journal)
TWORKOSKI, T., DAW, T., GLENN, D.M. "EFFECTS OF A ROOT BARRIER AND LOCALIZED FERTILIZER APPLICATION ON ROOT GROWTH OF YOUNG PEACH (PRUNUS PERSICA)TREES.". PLANT GROWTH REGULATOR SOCIETY OF AMERICA QUARTERLY. 2004. 31:133-142
Morphological and Hormonal Relationships in Shoots of Pillar and Standard Peach Trees - (Abstract Only)
Tworkoski, T. 2003. Morphological and hormonal relationships in shoots of pillar and standard peach trees. Society of America Plant Growth Regulator Proceedings. 30:190
Soil Amendment with Hydrophobic Kaolin Particles Reduce Weeds - (Abstract Only)
TAKEDA, F., GLENN, D.M., TWORKOSKI, T. SOIL AMENDMENT WITH HYDROPHOBIC KAOLIN PARTICLES REDUCE WEEDS. HORTSCIENCE. 2003. 38:659
N and P Uptake by Strawberry Plants Grown with Composted Poultry Litter - (Peer Reviewed Journal)
TWORKOSKI, T., PREUSCH, P., TAKEDA, F. N AND P UPTAKE BY STRAWBERRY PLANTS GROWN WITH COMPOSTED POULTRY LITTER. 2003. doi:10.1016/j.scienta.2003.12.005; Scientia Horticulturae 102 (2004) 91-103.
Regulating Vegetative Growth in Deciduous Fruit Trees - (Peer Reviewed Journal)
Miller, S.S., Tworkoski, T. 2003. Regulating vegetative growth in deciduous fruit trees. Plant Growth Regulation Society of America Quarterly.Volume 31, No. 1, Pgs 8 - 46.
Nitrogen and Phosphorus Availability and Weed Suppression from Composted Poultry Litter Applied As a Mulch in a Peach Orchard - (Peer Reviewed Journal) - (10-Jan-03)
ARS Research on Natural Products for Pest Management - (Peer Reviewed Journal)
AKEY, D.H., ARTHUR, F.H., DE LUCCA II, A.J., GIBSON, D.M., HARRISON JR, H.F., PETERSON, J.K., GEALY, D.R., TWORKOSKI, T., WILSON, C.L., MORRIS, J.B. ARS RESEARCH ON NATURAL PRODUCTS FOR PEST MANAGEMENT. PEST MANAGEMENT SCIENCE. 2003. v. 59. p. 708-717.
Weed Suppression with Organic Mulch in Orchards - (Proceedings) - (13-Dec-02)
Apple and Peach Tree Response to Scoring and Restriction of Trunk - (Proceedings) - (03-Oct-02)
The Synthesis and Biological Evaluation of Eugenol Derivatives As Potential Herbicidal Agents - (Proceedings) - (15-Sep-02)
Nitrogen and Phosphorus Availability and Plant Uptake from Soil Treated with Composted and Uncomposted Poultry Litter. - (Proceedings)
TWORKOSKI, T., ADLER, P.R., SIKORA, L., TAKEDA, F., FERRIER, D., PREUSCH, P. NITROGEN AND PHOSPHORUS AVAILABILITY AND PLANT UPTAKE FROM SOIL TREATED WITH COMPOSTED AND UNCOMPOSTED POULTRY LITTER. Proceedings of the Composting in the Southeast Conf and Exp Workshop. 2002. pp 1-9.
Two-Year Response of Apple and Peach Trees to Scoring and Restriction of Trunks. - (Proceedings) - (11-Jul-02)
Nitrogen Mineralization Rates and Phosphorus Availability in Composted and Uncomposted Poultry Litter - (Peer Reviewed Journal)
Preusch, P.L., Adler, P.R., Sikora, L.J., Tworkoski, T. 2002. Nitrogen mineralization rates and phosphorus availability in composted and uncomposted poultry litter. Journal of Environmental Quality. 31:2051-2057.
Herbicide Effects of Essential Oils - (Peer Reviewed Journal) - (01-May-02)
Bioactive Properties of Eugenol and Its Synthetic Deviations - (Abstract Only) - (26-Apr-02)
Herbicide Effects of Essential Oils. - (Abstract Only) - (21-Sep-01)
Response of Bean to Particle Film Applications - (Peer Reviewed Journal) - (09-Sep-01)
Root and Shoot Characteristics of Peach Tree with Different Growth Habits - (Peer Reviewed Journal) - (07-Aug-01)
Diversity and Biological Control Potential of M.PULCHERRIMA Strains Againstblue Mold of Apple - (Peer Reviewed Journal) - (01-Aug-01)
Characterizing the Mechanism of Biological Control of Postharvest Diseases on Fruits with a Simple Method to Study Competition for Nutrients - (Peer Reviewed Journal) - (31-Jul-01)
Herbicide Effects of Essential Oils - (Abstract Only) - (22-Jun-01)
Effects of Residues of Diuron, Simazine, and Terbacil on Newly Planted Apple and Peach Trees - (Peer Reviewed Journal) - (19-Jun-01)
Response of Mature Peach Trees to Grass Competition - (Peer Reviewed Journal) - (21-May-01)
Effects of Long Term Herbicide Use on the Orchard Floor and Growth of Young Fruit Trees - (Proceedings) - (30-Mar-01)
Phenotypic, Genetic and Functional Characterizing of Metchnikowia Pulcherrima Strains That Control Blue Mold of Apple - (Abstract Only) - (15-Mar-01)
Yield of Peach Trees Grown with Different Competition and Pruning Intensities - (Abstract Only) - (01-Feb-01)
Apple and Peach Tree Response to Scoring and Restriction of Trunks - (Proceedings) - (25-Nov-00)
Root Characteristics of Peach Trees with Different Short Growth Habits - (Proceedings) - (15-Oct-00)
The Effect of Nitrogen on Stolon and Ramet Growth in Four Genotypes of Fragaria Chiloensis L. - (Peer Reviewed Journal) - (26-Jun-00)
Root Characteristics of Peach Trees with Different Shoot Growth Habits - (Proceedings) - (21-Feb-00)
Weed Suppression and N and P Minieralization in An Orchard Mulched with Composted Poultry Litter - (Proceedings) - (20-Feb-00)
Weed Control and Mineralization of Nitrogen and Phospohorus from Composted Poultry Mulch in a West Virginia Peach Orchard - (Abstract Only) - (18-Jan-00)
Soil Residues Following Repeat Applications of Diuron, Simazine, and Terbacil - (Peer Reviewed Journal) - (29-Nov-99)
Weed Community Changes Following Applications of Diuron, Simazine and Terbacil - (Peer Reviewed Journal) - (26-Nov-99)
Effects of Residues of Diuron, Simazine, and Terbacil on Newly Planted Apple and Peach Trees - (Proceedings) - (26-Nov-99)
Root Restriction and Fertilizer Effects on Young Peach Trees - (Abstract Only) - (24-Nov-99)
Response of Mature Peach Trees to Grass Competition - (Proceedings) - (22-Nov-99)
Nitrogen Minerlization and Phosphorus Availability of Composted and Uncomposted Poultry Litter - (Abstract Only)
Tworkoski, T., Preusch, P.L., Adler, P.R., Sikora, L.J., Takeda, F., Ferrier, D. 1999. Nitrogen minerlization and phosphorus availability of composted and uncomposted poultry litter. Agronomy Abstracts. p. 37.
Nitrogen Mineralization and Phosphorus Availability of Composted and Uncomposted Poultry Litter - (Proceedings)
Preusch, P.L., Adler, P.R., Sikora, L.J., Takeda, F., Ferrier, D., Tworkoski, T. 1999. Nitrogen mineralization and phosphorus availability of composted and uncomposted poultry litter. [abstract]. Proceedings of the Soil Science Society of America Annual Meeting; Salt Lake City, UT, November 1999. Div S-8, p. 314.
Last Modified: 2/23/2010
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